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Starzomski, Brian M.; Bondrup-Nielsen, Soren . (2002). Analysis of movement and the consequence for metapopulation structure of the forked fungus beetle, Bolitotherus cornutus Panzer (Tenebrionidae). Écoscience, 9(1), 20–27. doi:10.1080/11956860.2002.11682686 
10.1080/11956860.2002.11682686